MARION, Ala. -- A former state trooper surrendered Thursday on a murder charge in the 1965 shooting death of a black man during a civil rights protest, a killing that led to the ''Bloody Sunday'' march and the passage of the Voting Rights Act.

Former trooper James Bonard Fowler, who contends he fired in self-defense in a struggle over a gun, was charged with first-degree and second-degree murder in the fatal shooting of 26-year-old Jimmie Lee Jackson.