The 16th-Century paintings were returned to the heirs of their former owners, Jakob and Rosa Oppenheimer, who died during the Holocaust.

After World War II, the paintings ended up in the collection of US newspaper tycoon, William Randolph Hearst.

They have been on display at a California museum for over 30 years.

Copies of two of the paintings will stay on display at Hearst Castle along with the original of the third painting belonging to the couple...